The Uncertainty of Gene Expression Networks

When he's saying uncertainty, he means the inability to perfectly predict the future. He goes on to say that ignorance of fundamental laws does not tell you that a messy looking pattern really is messy. But because we've already found the master order, we can guess that we won't find any additional secret patterns that will make biology as easy as a sequence of cubes. The elegance, the rules that govern physics is too far down to help us overcome our lexical and logical uncertainty.