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Behavioural Ecology

In Our Time

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What Are Tinbergen's Four Questions?

Rebecca Kellen: Tinbergen's four questions were essential in moving the study of animal behaviour beyond anecdotal evidence. She says he set out these four questions which are four kinds of ways that we can start to begin to investigate animal behaviour. The question really is, for what purpose has this behaviour evolved? How does it function to promote an animal's fitness? she asks.

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