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The Sleeping Beauty of Bacteria
Many of the enzymes that are running these metabolic networks are unexpectedly proboscuous in the sense that they don't just do one reaction, right? They have this capacity to also do other things, maybe not as well or as rapidly, but that those other things represent a kind of capacitance for rapid evolutionary change. So even if there were no promiscuous enzymes, that is, every enzyme can only catalyze a single reaction. This property of metabolism would still exist.