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What makes ant colonies robust?

Simplifying Complexity

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Redundancy Is an Emergent Property of the System

Redundancy alone is simply not enough. Redundancy does play a role, but it's not the only thing that plays a role. The fact that I can scoop up a thousand ants from a colony doesn't hurt the colony too much because there are at least another ten thousand that could take that place. And if there had been more ants, then the system would have been more robust.

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