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The Big Picture
Fitness is hard to quantify in a reliable way that every one would agree onye. The idea of kin selection and inclusive fitness was introduced by william bill hamilton, who published two papers in the 19 sixties. Hamilton realized that you shouldn't just think about for organisms, the offspring that a specific individual has, but should also account for other offspring that this focal individual is corsally responsible for. Fitness scaled by the relatedness that you have to those individuals, in a way that if your brother has it, that counts more than if your cousin does it, and so on. That then can be maximized under certain circumstances - even though they do not personally have any